Late Pliocene to Early Pleistocene; Extinct.
Southern Florida.
Original Description (from Mansfield, 1932, p. 48):
"Test large, suborbicular, and moderately high; upper surface convex and broadly rounded, the posterior surface more gently inclined than the anterior; lower surface nearly flat except in the area surrounding the peristome, where it is shallowly concave. Apical system, situated opposite the peristome, is rather large, granular, and slightly elevated; so far as revealed, a genital pore is at the juncture of the petals and a smaller radial pore is opposite each petal. Ambulacral areas petaloid at dorsal portions. Petals rather long, extending nearly to the ambitus, expanding to about one-third their length from the apical system, then gradually contracting distally, and nearly closing at their extremities; poriferous zones rather wide, shallowly depressed; pores nearly equal in size and rounded in outline; pairs of pores conjugate. Interporiferous areas weakly tumid. Posterior interambulacrum weakly medially arched. Periproct rather large, longest transversely; supramarginal, the lower margin being about 4 millimeters above the ambitus; the upper arched margin slightly overhangs the aperture. Peristome excentric anteriorly, pentagonal, transversely elongate, and surrounded by a large well-defined floscelle with prominent bourrelets. The outer pores of the floscelle are more direct and more regularly placed; the inner ones are more irregularly placed and some of them are arranged in two rows. The surface of the test is closely set with scrobiculate tubercles.
Dimensions: Cotypes (catalogue No. 371329, U. S. Nat. Mus.): Larger slightly crushed cotype with preserved periproct, length, 73 millimeters; width, 74 millimeters; height, 29 millimeters. Smaller cotype (posterior end broken off), width, 57 millimeters; height, 32 millimeters. Paratype (catalogue No. 371330, U. S. Nat. Mus.), length, 75 millimeters (posterior end broken) ; width, 64 millimeters; height, 35 millimeters.
Occurrence: Station 1/1177, Tamiami Trail, 5 miles east of Carnestown and about 7 miles northeast of Everglades, Collier County, Fla. (type locality); station 1/1179, Tamiami Trail, 9 miles west of Pinecrest (sec. 13, T. 54 S., R. 32 E.), Monroe County, Fla. (paratype locality). Near Moore Haven, Glades County, Fla.; probably dredged from the canal. Only one fairly well preserved specimen, which was embedded in limestone, has been collected near Moore Haven; it is in the collection of the Florida Geological Survey.
Cassidulus evergladensis resembles, in a general way, the figured type of Cassidulus (Pygorhynchus) alabamensis Twitchell, a species collected at the Natural Bridge, St. Stephens, Washington County, Ala., but the lower surface of the test of that species is more concave longitudinally, and the periproct is at a greater distance above the ambitus. The same authors refer the "St. Stephens limestone" (upper part) to the lower Oligocene."
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